Scottish Highlands

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0:00:02 > 0:00:05Our planet is the greatest living puzzle in the universe.

0:00:05 > 0:00:09A collection of worlds within worlds,

0:00:09 > 0:00:13each one a self-contained ecosystem bursting with life.

0:00:17 > 0:00:19But how do they work?

0:00:21 > 0:00:23The intricate web of relationships

0:00:23 > 0:00:26and the influence of natural forces

0:00:26 > 0:00:31makes each microworld complex and unique.

0:00:33 > 0:00:35So to discover their secrets,

0:00:35 > 0:00:38we need to explore them one by one.

0:00:40 > 0:00:43Untangle their interlocking pieces

0:00:43 > 0:00:47and ultimately reveal the vital piece,

0:00:47 > 0:00:49the key to life itself,

0:00:49 > 0:00:53hidden deep within each of nature's microworlds.

0:01:06 > 0:01:08The Scottish Highlands.

0:01:08 > 0:01:11An ancient landscape of startling beauty.

0:01:14 > 0:01:16Craggy peaks peer over a terrain

0:01:16 > 0:01:19of thick forest and wide open space.

0:01:21 > 0:01:26Two contrasting habitats united by their battle with the elements.

0:01:28 > 0:01:32Presided over by Scottish emblems

0:01:32 > 0:01:34like golden eagles

0:01:34 > 0:01:37and red stags.

0:01:37 > 0:01:40And it's home to some of the rarest animals in Britain.

0:01:43 > 0:01:44The hand of man has been felt

0:01:44 > 0:01:46throughout the history of the Highlands,

0:01:46 > 0:01:49and yet its wildlife remains iconic.

0:01:55 > 0:01:58How have the Highlands remained a home for these species

0:01:58 > 0:02:00despite the pressure of the modern world?

0:02:02 > 0:02:08To find out, let's explore its deep forest and open heather moors,

0:02:08 > 0:02:11discover the impact of the elements on these landscapes,

0:02:11 > 0:02:14and the role of their key characters

0:02:14 > 0:02:18in keeping this remote corner of the British Isles wild.

0:02:26 > 0:02:29There's a famous saying about Scotland -

0:02:29 > 0:02:32it doesn't have a climate, just weather.

0:02:37 > 0:02:41Nowhere is this more true than in the Highlands.

0:02:43 > 0:02:48It's a landscape at the mercy of gale-force Atlantic winds.

0:02:50 > 0:02:52Winters have been known to hit -20.

0:02:53 > 0:02:56And when it rains, it pours.

0:02:58 > 0:03:02Have the elements had a hand in keeping this place wild?

0:03:03 > 0:03:07Up to five metres of rain fall here every year -

0:03:07 > 0:03:10that's the same as in some rainforests.

0:03:10 > 0:03:12With most other parts of Britain

0:03:12 > 0:03:14receiving less than a quarter of this,

0:03:14 > 0:03:17it's easy to see why humans might have chosen

0:03:17 > 0:03:19to set up shop elsewhere.

0:03:20 > 0:03:22But the Highlands' wild residents

0:03:22 > 0:03:24are made of sterner stuff.

0:03:26 > 0:03:29In fact, some rely on rain.

0:03:31 > 0:03:34From the soggy earth emerges a creature

0:03:34 > 0:03:36feared throughout the Highlands.

0:03:38 > 0:03:41The scourge of Scotland -

0:03:41 > 0:03:43the midge.

0:03:44 > 0:03:47This tiny millimetre-long vampire

0:03:47 > 0:03:50hatches from eggs laid in the rain-soaked ground.

0:03:55 > 0:03:57As the midge feeds,

0:03:57 > 0:03:59it sends off a chemical signal

0:03:59 > 0:04:02inviting others in the area to join the feast.

0:04:04 > 0:04:07Cutting the skin with their scissor-like mouthparts

0:04:07 > 0:04:09and sucking up the pooling blood.

0:04:14 > 0:04:18And with an estimated 40,000 capable of landing on a deer

0:04:18 > 0:04:21or similar-sized animal within an hour,

0:04:21 > 0:04:23they can make life unbearable.

0:04:25 > 0:04:27Midges die off before winter,

0:04:27 > 0:04:29but the change in seasons

0:04:29 > 0:04:32doesn't make life in the Highlands any easier,

0:04:32 > 0:04:35as the rain turns to snow.

0:04:37 > 0:04:41On cue, the mountain hare population turns from russet brown

0:04:41 > 0:04:43to brilliant white.

0:04:46 > 0:04:49The new coat is thicker and warmer,

0:04:49 > 0:04:52but it also helps to keep the hare safe from predators.

0:04:53 > 0:04:56A brown hare on a white background

0:04:56 > 0:04:58would stick out like a sore thumb.

0:05:05 > 0:05:08The ptarmigan employs the same colour-changing tactics.

0:05:10 > 0:05:13Its feathery feet increase the surface area,

0:05:13 > 0:05:14acting like snow shoes,

0:05:14 > 0:05:18preventing the ptarmigan from sinking into the deep snow.

0:05:19 > 0:05:23Humans would struggle to live in these cold conditions.

0:05:23 > 0:05:27But surprisingly, these hardy animals can only survive here

0:05:27 > 0:05:30because of the Highland's extreme weather.

0:05:32 > 0:05:35As Britain melted from its most recent ice age,

0:05:35 > 0:05:37around 12,000 years ago,

0:05:37 > 0:05:41animals adapted for arctic conditions were forced back

0:05:41 > 0:05:45to the very highest, coldest points of the British Isles.

0:05:46 > 0:05:51While humans busily colonised the rest of the newly temperate UK,

0:05:51 > 0:05:53the Highlands provided a climate

0:05:53 > 0:05:56in which these animal refugees could survive.

0:06:01 > 0:06:05So, has the tough climate kept the Highlands wild?

0:06:06 > 0:06:08While it's true that, as a rule,

0:06:08 > 0:06:11the highest, coldest, wettest parts of the world

0:06:11 > 0:06:13are some of the least developed,

0:06:13 > 0:06:16actually, the Highlands share a similar climate

0:06:16 > 0:06:19with major cities in other parts of the world.

0:06:19 > 0:06:22So, if weather alone does not keep a place wild,

0:06:22 > 0:06:26the Highlands must have another secret up their sleeve.

0:06:28 > 0:06:31There's less than 50 people per square mile here

0:06:31 > 0:06:34compared with 600 in the rest of the UK.

0:06:35 > 0:06:38Perhaps the landscape has a part to play.

0:06:42 > 0:06:45There's one very special, ancient habitat

0:06:45 > 0:06:48that's found nowhere else in the United Kingdom.

0:06:54 > 0:06:56The Caledonian forest.

0:06:56 > 0:06:59Dominated by the gigantic Scots pine.

0:07:01 > 0:07:04Growing up to 20 metres high

0:07:04 > 0:07:06and a solid three metres in girth.

0:07:08 > 0:07:12Each tree can live for a very respectable 250 years...

0:07:15 > 0:07:19..during which time, it supports an array of Scottish wildlife.

0:07:20 > 0:07:24It's the backbone of the Caledonian community.

0:07:27 > 0:07:32A familiar character is on the hunt for food among its branches.

0:07:32 > 0:07:35The red squirrel's agility allows it to reach

0:07:35 > 0:07:39the very top branches of the pine,

0:07:39 > 0:07:41home to some of the choicest cones.

0:07:46 > 0:07:50It's been estimated that a single squirrel can eat the seeds

0:07:50 > 0:07:53from as many as 20,000 pine cones in a year.

0:07:59 > 0:08:01Squirrels have the sharp incisors

0:08:01 > 0:08:03common with the rest of the rodent family,

0:08:03 > 0:08:06which they use to tear the tough cone apart.

0:08:08 > 0:08:10Perfect tools for the job.

0:08:10 > 0:08:12But even so, these teeth will need to grow

0:08:12 > 0:08:15as much as 15 centimetres a year

0:08:15 > 0:08:17to keep up with wear and tear.

0:08:23 > 0:08:27The squirrel knows just how tough the winters can be,

0:08:27 > 0:08:30so it plans ahead,

0:08:30 > 0:08:32burying stores for leaner times.

0:08:36 > 0:08:38Scent glands in its cheeks help

0:08:38 > 0:08:41to guide it back to the cache.

0:08:41 > 0:08:43But it might be months

0:08:43 > 0:08:46before the squirrel needs to excavate its hoard

0:08:46 > 0:08:50and, unsurprisingly, some seeds are forgotten

0:08:50 > 0:08:52or dropped by messy eating.

0:08:56 > 0:08:58On the forest floor,

0:08:58 > 0:09:00they have a chance of germinating,

0:09:00 > 0:09:02bolstering the pine population.

0:09:02 > 0:09:05It's a mutually beneficial relationship

0:09:05 > 0:09:07that's been going strong for millennia.

0:09:11 > 0:09:15Food is not the only reason the squirrel prefers the Scots pine.

0:09:16 > 0:09:20The bark blends perfectly with its rusty red fur

0:09:20 > 0:09:22and camouflage is all important

0:09:22 > 0:09:26when you have a predator that's equally at home in the branches.

0:09:32 > 0:09:35The pine marten rivals the squirrel's agility...

0:09:38 > 0:09:40..and in a high tree top chase,

0:09:40 > 0:09:42is one of few predators able to catch it.

0:09:54 > 0:09:57A smaller male wants in on the meal,

0:09:57 > 0:09:59but he's testing his luck.

0:10:02 > 0:10:05This pine marten is in no mood for sharing...

0:10:05 > 0:10:07IT SQUEALS

0:10:07 > 0:10:10..and chases the chancer out to a thin branch.

0:10:10 > 0:10:13IT SQUEALS

0:10:15 > 0:10:17The squirrel makes a good meal

0:10:17 > 0:10:20in an environment where every calorie counts.

0:10:20 > 0:10:24The young male will just have to find his own food.

0:10:27 > 0:10:28And by the looks of it,

0:10:28 > 0:10:30he could be going hungry.

0:10:36 > 0:10:38Pine martens are agile enough

0:10:38 > 0:10:41to also prey on the pine's resident birds.

0:10:43 > 0:10:46Like the Scottish crossbill,

0:10:46 > 0:10:49an endemic species found in Highland Caledonian forests.

0:10:54 > 0:10:58The bird's shape reveals its close relationship with the pine.

0:10:59 > 0:11:02Its curved, slightly crossed beak is perfectly adapted

0:11:02 > 0:11:05to prise seeds from tough cones.

0:11:09 > 0:11:11It eats little else.

0:11:16 > 0:11:18The tiny crested tit,

0:11:18 > 0:11:19one of Britain's rarest birds,

0:11:19 > 0:11:22is equally at home in the Caledonian forest.

0:11:24 > 0:11:26The tit supports the pine

0:11:26 > 0:11:29by winkling insects from crevasses in the bark

0:11:29 > 0:11:32and preventing burrowing larvae from doing serious damage.

0:11:34 > 0:11:38But birds are not the only species that groom the tree.

0:11:40 > 0:11:44The caterpillars of the pine looper moth and sawfly

0:11:44 > 0:11:46mimic the needles that they feed on.

0:11:46 > 0:11:51These caterpillars can defoliate a whole tree.

0:11:52 > 0:11:54But the Scots pine has an ally.

0:11:59 > 0:12:01Wood ants, on patrol.

0:12:06 > 0:12:09They detect their prey by vibration,

0:12:09 > 0:12:11stalking and attacking.

0:12:19 > 0:12:21The larvae may be twice the size of the ant,

0:12:21 > 0:12:23but this poses little problem,

0:12:23 > 0:12:26because these ants are team players...

0:12:28 > 0:12:32..dragging the larvae from the needles back to their nest.

0:12:33 > 0:12:37The largest ant in Britain has a suitably impressive home.

0:12:37 > 0:12:43Each metre-high mound holds up to 100,000 ants at a time.

0:12:44 > 0:12:48The nest is constructed almost entirely from pine needles.

0:12:53 > 0:12:56With a queen ant living for anything up to 15 years,

0:12:56 > 0:13:01the nest must be constantly repaired and improved to see out her reign.

0:13:02 > 0:13:05Each one is a feat of engineering.

0:13:10 > 0:13:11With the Scots pine providing

0:13:11 > 0:13:14both food and construction material for the wood ant,

0:13:14 > 0:13:17it's no wonder that their nests are a common site

0:13:17 > 0:13:19throughout the Caledonian forest.

0:13:23 > 0:13:25In fact, so many of the forest's species

0:13:25 > 0:13:28are interconnected with the Scots pine,

0:13:28 > 0:13:30relying on it for a food source,

0:13:30 > 0:13:32a nursery or a home,

0:13:32 > 0:13:36that it's known as a keystone species.

0:13:36 > 0:13:39The backbone of the forest community

0:13:39 > 0:13:42and where many trees grow together,

0:13:42 > 0:13:46the forest habitat supports larger species.

0:13:46 > 0:13:48CLICKING SOUND

0:13:50 > 0:13:52IT CLICKS

0:13:52 > 0:13:56This strange call belongs to the capercaillie,

0:13:56 > 0:13:59the largest member of the grouse family.

0:14:00 > 0:14:03The turkey-sized bird lives across much of Europe,

0:14:03 > 0:14:07but, in Britain, is predominantly found in the Highlands.

0:14:07 > 0:14:08IT CLICKS

0:14:08 > 0:14:11This is actually a mating call.

0:14:11 > 0:14:13It might not sound particularly enticing,

0:14:13 > 0:14:18but it's widely believed that the subtleties of the love song

0:14:18 > 0:14:20are too low for human ears to register.

0:14:21 > 0:14:23Capercaillies have been breeding in the Highlands

0:14:23 > 0:14:26since the ice age gave way to forest.

0:14:33 > 0:14:36Stands of Scots pine set a stage for their mating displays

0:14:36 > 0:14:40from the Highlands to Western Asia

0:14:40 > 0:14:43and the males are dressed to impress.

0:14:49 > 0:14:53With the affection of the more drably coloured hens at stake...

0:14:56 > 0:15:00..testosterone-fuelled tussles regularly break out.

0:15:00 > 0:15:01CLICKING

0:15:18 > 0:15:21The chance of injury is very real.

0:15:25 > 0:15:27The victor gets his girl

0:15:27 > 0:15:29and the chance to mate.

0:15:29 > 0:15:32IT CLICKS

0:15:35 > 0:15:38The Caledonian forests and their Scots pine

0:15:38 > 0:15:41are synonymous with the Highland wilderness.

0:15:43 > 0:15:47But they're only one habitat within this microworld.

0:15:49 > 0:15:51The forest exists side by side

0:15:51 > 0:15:54with another completely contrasting landscape...

0:15:56 > 0:15:59..moorland, as open as the forest is dense.

0:16:07 > 0:16:09Scotland's largest living mammal,

0:16:09 > 0:16:10the red deer,

0:16:10 > 0:16:12needs both these habitats.

0:16:15 > 0:16:18Deer feed on new pine shoots,

0:16:18 > 0:16:21but need open space to breed.

0:16:21 > 0:16:25IT BELLOWS

0:16:30 > 0:16:33Large males bellow across the heather,

0:16:33 > 0:16:36advertising their superiority.

0:16:38 > 0:16:41The stag rounds up a herd of females.

0:16:41 > 0:16:43By keeping them under his watch,

0:16:43 > 0:16:45he can ensure that no other male

0:16:45 > 0:16:48breeds with them and that only his genes are passed on.

0:16:51 > 0:16:54But that's easier said than done.

0:16:54 > 0:16:57Other males will want the females for themselves

0:16:57 > 0:17:00and are ready to fight for the right.

0:17:00 > 0:17:03THEY BELLOW

0:17:09 > 0:17:12These are ferocious battles

0:17:12 > 0:17:13to prove strength

0:17:13 > 0:17:14and determine dominance.

0:17:23 > 0:17:26The rut has been fought in the open heather moorlands

0:17:26 > 0:17:29for more than 11,000 years.

0:17:29 > 0:17:34It's the perfect arena, stretching for miles without a single tree,

0:17:34 > 0:17:37let alone sign of human habitation.

0:17:40 > 0:17:42Without natural shelter,

0:17:42 > 0:17:45it's no wonder humans avoided the moors,

0:17:45 > 0:17:48but deer are not the only animals to thrive.

0:17:49 > 0:17:52Despite first appearances,

0:17:52 > 0:17:56the open moorland is as rich with life as the pine forest.

0:17:59 > 0:18:02The red grouse is as dependent on the heather

0:18:02 > 0:18:05as its cousin, the capercaillie, is on the pine.

0:18:05 > 0:18:07IT CLUCKS

0:18:07 > 0:18:11Feeding on seeds, flowers and shoots as the seasons change.

0:18:12 > 0:18:14The heather, like the Scots pine,

0:18:14 > 0:18:17relies on the relationships with other species.

0:18:19 > 0:18:22Its striking purple flowers are pollinated by bees,

0:18:22 > 0:18:24while passing animals help

0:18:24 > 0:18:26to distribute its seeds.

0:18:30 > 0:18:32And it creates a territory

0:18:32 > 0:18:36for one of the most powerful birds on Earth.

0:18:37 > 0:18:39The golden eagle,

0:18:39 > 0:18:42a symbol of the wild Highlands.

0:18:42 > 0:18:45It's a predator, adapted for purpose.

0:18:47 > 0:18:51Keen eyesight allows it to scour the heather for movement

0:18:51 > 0:18:53from up to two kilometres away.

0:18:56 > 0:18:59Powerful talons and beak help it to dispatch prey

0:18:59 > 0:19:01many times its body weight.

0:19:04 > 0:19:08And this formidable bird needs these open spaces...

0:19:09 > 0:19:11..to hunt.

0:19:20 > 0:19:24Following a particularly cold winter, the snow's melted,

0:19:24 > 0:19:27but the mountain hare's fur hasn't quite caught up.

0:19:29 > 0:19:31The white that provided camouflage a month before

0:19:31 > 0:19:34is now more of an advertisement.

0:19:38 > 0:19:39With its two metre wing span,

0:19:39 > 0:19:41the eagle soars on wind currents,

0:19:41 > 0:19:44constantly spotting for movement below.

0:19:47 > 0:19:52While the hare is unaware of the danger overhead,

0:19:52 > 0:19:54the eagle locks in.

0:19:56 > 0:19:58But the hare is no sitting target.

0:19:58 > 0:19:59Once it spots the threat,

0:19:59 > 0:20:01it will sprint through the heather.

0:20:02 > 0:20:05By using pre-practiced routes that it knows are obstacle free,

0:20:05 > 0:20:09the hare is capable of hitting 60 kilometres an hour.

0:20:09 > 0:20:11But with the eagle topping out

0:20:11 > 0:20:13at three times this,

0:20:13 > 0:20:15it needs something more than speed to survive.

0:20:23 > 0:20:25So the hare dodges and weaves,

0:20:25 > 0:20:28throwing itself out of the eagle's trajectory.

0:20:32 > 0:20:36A last-minute zigzag saves the hare by a whisker.

0:20:44 > 0:20:47Heather plants can grow for around 30 years.

0:20:48 > 0:20:51After this time, they degenerate and die back.

0:20:51 > 0:20:55Bacteria in the soil will break them down.

0:20:55 > 0:20:58The increased nutrients that this breakdown provides

0:20:58 > 0:21:03is just enough for larger, hardy plants to begin to grow.

0:21:03 > 0:21:05Over time, heather will be replaced

0:21:05 > 0:21:10by trees and, eventually, forest.

0:21:10 > 0:21:13But something doesn't quite make sense.

0:21:13 > 0:21:17If this natural succession was taking place,

0:21:17 > 0:21:20the Highlands should be full of trees.

0:21:22 > 0:21:27Instead, Caledonian forest only exists in small pockets.

0:21:30 > 0:21:33This explains why some of the species that it supports

0:21:33 > 0:21:35are so rare in Britain,

0:21:35 > 0:21:38as there's just not enough habitat to go around.

0:21:40 > 0:21:42But it hasn't always been this way.

0:21:43 > 0:21:46Some 10,000 years ago,

0:21:46 > 0:21:49after the end of the last ice age,

0:21:49 > 0:21:51Scotland was covered in trees.

0:21:54 > 0:21:57These primordial forests would have been rich with life,

0:21:57 > 0:22:01wolves, bears, wild boar,

0:22:01 > 0:22:03animals missing from Scotland today.

0:22:03 > 0:22:04WOLF HOWLS

0:22:04 > 0:22:08So what has happened to the Highlands?

0:22:08 > 0:22:10How does a forest just disappear?

0:22:11 > 0:22:16What's allowed the heather to take hold in such vast areas?

0:22:16 > 0:22:20And how has the natural succession of the trees been halted?

0:22:24 > 0:22:29The soil below the heather reveals the answer to this conundrum.

0:22:30 > 0:22:34By studying remnants of the habitats from thousands of years back,

0:22:34 > 0:22:36scientists have been able to determine

0:22:36 > 0:22:41that there was still forest in the Highlands up to 5,000 years ago.

0:22:42 > 0:22:43Before disaster struck.

0:22:47 > 0:22:49The climate changed.

0:22:51 > 0:22:55A small fluctuation compared with the chill of the ice age,

0:22:55 > 0:22:59but enough to make the Highlands wetter.

0:22:59 > 0:23:01The deluge that followed was far more serious

0:23:01 > 0:23:04than the worst of modern-day Scottish winters.

0:23:06 > 0:23:08The forests literally drowned.

0:23:11 > 0:23:12It's thought that three quarters

0:23:12 > 0:23:15of the original Caledonian forest rotted away,

0:23:15 > 0:23:18creating vast bare spaces.

0:23:20 > 0:23:25Which suited not only the tough heather, but another species.

0:23:27 > 0:23:28Humans.

0:23:30 > 0:23:34Open land allowed communities to farm crops, build homes

0:23:34 > 0:23:36and raise livestock.

0:23:38 > 0:23:42For the first time, humans became a dominant Highland species.

0:23:44 > 0:23:46The climate eventually settled,

0:23:46 > 0:23:51but only a tiny 10% of the once vast Caledonian forest remained.

0:23:53 > 0:23:57The landscape that we recognise as the Scottish Highlands today

0:23:57 > 0:23:59had been created.

0:23:59 > 0:24:04This was, of course, great news for open-space-loving species

0:24:04 > 0:24:06like the grouse and mountain hare.

0:24:08 > 0:24:11IT HOWLS

0:24:11 > 0:24:12But many of the animals

0:24:12 > 0:24:14that relied on the forest -

0:24:14 > 0:24:15wolves,

0:24:15 > 0:24:17the capercaillie,

0:24:17 > 0:24:20red squirrel

0:24:20 > 0:24:22and bears became extinct.

0:24:25 > 0:24:28Those species that could survive were excluded

0:24:28 > 0:24:30to small pockets of woodland.

0:24:32 > 0:24:36Over the centuries, the land use of the Highlands shifted.

0:24:36 > 0:24:40Farms were cleared to make way for deer and grouse hunting.

0:24:40 > 0:24:43The Highlands were kept bare.

0:24:43 > 0:24:46Trophy species relied on healthy heather,

0:24:46 > 0:24:49and so patches were burnt to encourage new growth

0:24:49 > 0:24:52and extend the plants natural 30-year cycle.

0:24:58 > 0:25:02A new habitat was created,

0:25:02 > 0:25:06a patchwork of old and young heather growing side by side,

0:25:06 > 0:25:11in which not only the grouse, but all moorland species could flourish.

0:25:13 > 0:25:16But this wasn't a natural habitat.

0:25:19 > 0:25:23Man was artificially managing the Highland's wild populations.

0:25:26 > 0:25:30Humans had become an intrinsic part of the ecosystem.

0:25:34 > 0:25:36Scotland's heather moorlands were thriving,

0:25:36 > 0:25:40but the few remaining areas of forest and their inhabitants

0:25:40 > 0:25:41were still isolated.

0:25:43 > 0:25:46It was an environmental disaster that triggered the demise

0:25:46 > 0:25:49of the Caledonian forest 5,000 years before,

0:25:49 > 0:25:53so it's fitting that another would start to bring it back.

0:25:54 > 0:25:57But this time it wasn't brought about by natural forces.

0:26:05 > 0:26:08Two world wars left Britain's natural resources ravaged.

0:26:12 > 0:26:13New trees needed planting

0:26:13 > 0:26:16and, although their primary purpose was for timber,

0:26:16 > 0:26:20these forests also encouraged species back into Scotland

0:26:20 > 0:26:22that hadn't been seen in decades.

0:26:24 > 0:26:26Red squirrels re-colonised,

0:26:26 > 0:26:30supporting greater numbers of their predators - pine martens.

0:26:33 > 0:26:38Pine-loving populations of the rare crested tit and the crossbill grew.

0:26:43 > 0:26:47Landowners began to take responsibility for the wildlife.

0:26:48 > 0:26:52The focus shifted to forest protection and regeneration.

0:26:53 > 0:26:55But this posed a dilemma.

0:26:55 > 0:26:57Left to its own devices,

0:26:57 > 0:27:00the forest would naturally replace the heather habitat

0:27:00 > 0:27:04that now supported many more species than just the grouse.

0:27:04 > 0:27:08The regeneration of one couldn't come at the expense of the other.

0:27:11 > 0:27:15For these two habitats to co-exist in the Highlands,

0:27:15 > 0:27:18then humans would need to be actively involved.

0:27:21 > 0:27:22They'd have to site manage.

0:27:27 > 0:27:30Today, the Scottish Highlands are celebrated

0:27:30 > 0:27:32as the last truly wild place in Britain.

0:27:33 > 0:27:37But it is a wilderness under supervision.

0:27:39 > 0:27:43Forests are being planted and the heather managed.

0:27:43 > 0:27:46Species have been re-introduced.

0:27:46 > 0:27:50It is a microworld under reconstruction.

0:27:50 > 0:27:53One in which humans are as fundamental to success

0:27:53 > 0:27:56as any of its other animals or plants.

0:27:57 > 0:28:02The Scottish Highlands are, at once, both wild and controlled.

0:28:07 > 0:28:12People have had to learn how to go about regenerating an ecosystem.

0:28:14 > 0:28:18A long and delicate process that's still not complete.

0:28:19 > 0:28:23But the hope is that, under careful guardianship,

0:28:23 > 0:28:27these iconic characters and ice age survivors

0:28:27 > 0:28:32will have a place in the Highland landscape for centuries to come.

0:28:56 > 0:28:59Subtitles by Red Bee Media Ltd