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behind thousands of pages of drawings and notes including a | :01:44. | :01:47. | |
series of startlingly accurate anatomical sketches that lay | :01:47. | :01:52. | |
undiscovered for hundreds of years. To coincide with this year 's | :01:52. | :01:54. | |
Edinburgh International Festival a new exhibition at the Queen 's | :01:54. | :01:58. | |
Gallery at the Palace of Holyrood shows his exquisite studies | :01:58. | :02:04. | |
alongside state-of-the-art modern medical imagery revealing just how | :02:04. | :02:07. | |
close this Renaissance genius got to the truth that lies beneath the | :02:07. | :02:15. | |
skin. He shines out as somebody who made | :02:15. | :02:19. | |
enormous strides in his field. Thinking as an engineer, trying to | :02:19. | :02:26. | |
understand the mechanism of the body. It is absolutely accurate. | :02:26. | :02:30. | |
find it quite poetic he has used movement to illustrate something | :02:30. | :02:33. | |
anatomical. It gives you a depth to the drawings which is technically | :02:33. | :02:43. | |
:02:43. | :02:47. | ||
Vinci as a painter, but for the majority of his life he was also a | :02:47. | :02:51. | |
scientist designing robots, studying the property of water, endeavouring | :02:51. | :02:55. | |
to understand the secrets of light, but the scientific field in which he | :02:55. | :03:03. | |
most excelled was that of human anatomy. He first began to research | :03:03. | :03:07. | |
the human body to help him keep his paintings as true to nature as | :03:07. | :03:14. | |
possible. But the project soon took on a life of its own, when he filled | :03:14. | :03:21. | |
hundreds of pages of his notebooks with detailed sketches. His aim was | :03:21. | :03:25. | |
always to publish an illustrated treaty on the human body. But | :03:26. | :03:34. | |
tragically he never did. Today all of his notebooks are | :03:34. | :03:40. | |
scattered right across the world. And perhaps surprisingly almost all | :03:40. | :03:43. | |
of the anatomical ones are amongst the Royal collection 's greatest | :03:43. | :03:49. | |
treasures at Windsor Castle. The weird thing about them is for | :03:49. | :03:52. | |
hundreds of years the fruits of his research were essentially lost. | :03:52. | :03:56. | |
There is a wonderful story that the drawings languished for a long time | :03:56. | :04:01. | |
in a royal bureau, until the wife of George II, Queen Caroline, chanced | :04:01. | :04:07. | |
upon them. The reasons why they ended up in the Royal collection are | :04:07. | :04:13. | |
slightly complicated. Towards the end of the 16th century the sculptor | :04:13. | :04:20. | |
Pompeo Leoni bought a load of his papers and carved up, sometimes | :04:20. | :04:24. | |
literally, into different albums, including one that consist of about | :04:24. | :04:31. | |
600 sheets. This was probably acquired by Charles II sometimes | :04:31. | :04:40. | |
after the Civil War. Curator Martin Clayton is allowing me a rare | :04:40. | :04:46. | |
glimpse of these delicate works here at Windsor Castle. Before they are | :04:46. | :04:49. | |
packed away and transported to the exhibition in Edinburgh. | :04:49. | :04:56. | |
Did people in his lifetime know about the anatomical drawings? | :04:56. | :05:00. | |
that it is not the same as understanding their content. -- but | :05:00. | :05:06. | |
it is not the same. Until anatomists came along in the 18th century, | :05:06. | :05:09. | |
nobody really understood the content of this material, they knew there | :05:09. | :05:14. | |
were impressive but they did not know why. There can be no other | :05:14. | :05:17. | |
scientist whose work was as profoundly insightful as his was, | :05:18. | :05:27. | |
that has had so little impact on his chosen field. Martin has selected | :05:27. | :05:31. | |
for different drawings from the netbooks to give an overview of his | :05:31. | :05:40. | |
anatomical career -- notebooks. The first one dates from 1489. What is | :05:40. | :05:45. | |
astonishing is how beautiful the presentation is, to take the front | :05:45. | :05:50. | |
of one half and juxstapose the two sides say you can see the depth | :05:50. | :06:00. | |
:06:00. | :06:00. | ||
interlacing to the searches features is a brilliant demonstration. | :06:00. | :06:07. | |
my batch I find it hard to get my head around the fact he drew this, | :06:07. | :06:12. | |
he did touch this paper. This is from the spring of 15 oh wait, he | :06:12. | :06:16. | |
describes observing the death of the centenarian performing a | :06:16. | :06:26. | |
:06:26. | :06:29. | ||
post-mortem. The old man, a few hours before his death. Why does he | :06:29. | :06:38. | |
use the writing? It was easier.If we fast forward two years we find | :06:38. | :06:46. | |
this kind of sheet. This is from 1510, 1511, a sequence of | :06:46. | :06:50. | |
illustrations, deeper and deeper as he takes away individual muscles. | :06:50. | :06:55. | |
The density of observation and the quality of presentation you see is a | :06:55. | :06:59. | |
different order compared with what went before. | :06:59. | :07:04. | |
This final image is one of the most famous drawings he produced. It is | :07:04. | :07:08. | |
one of the few sheets in which he uses colour, the red chalk leaps out | :07:08. | :07:13. | |
at you from the page. That red, the startling use of red, gives the | :07:13. | :07:20. | |
foetus a sense of life, that is possibly lacking in some of the | :07:20. | :07:29. | |
Perhaps I am feeling especially susceptible at the moment because I | :07:29. | :07:34. | |
am about to become a father for the first time but that red chalk | :07:34. | :07:38. | |
drawing of a foetus curled up in the womb, such a ravishing, heart | :07:38. | :07:41. | |
stopping thing, seems to wriggle before your eyes and every single | :07:41. | :07:47. | |
one of those sheet is similarly animated by the alacrity of his | :07:47. | :07:51. | |
thoughts because each one jostles and teams with observations and | :07:51. | :07:54. | |
ideas and reflections, a bit like the fossilised remains of somebody's | :07:54. | :08:04. | |
:08:04. | :08:12. | ||
synapses firing on all cylinders now be among the treasures at | :08:12. | :08:17. | |
Windsor Castle, but the first regatta work on them here in Milan. | :08:17. | :08:22. | |
-- but he first began work on them. He came to the city as a fully | :08:22. | :08:26. | |
fledged artist around 1482 and the years he spent in this part of Italy | :08:26. | :08:32. | |
would prove crucial. He arrived here to work at the Court | :08:32. | :08:35. | |
of Ludovico Sforza, the ruler of Milan whose nickname was Il Moro on | :08:35. | :08:45. | |
:08:45. | :08:46. | ||
account of his, Jim. -- his complexion. Leonardo arrived | :08:46. | :08:49. | |
brandishing a musical instrument which he had fashioned to resemble a | :08:49. | :08:56. | |
horsey skull. -- horse's goal. It was an advertisement for his | :08:56. | :09:02. | |
ingenuity in different talents. Working for Ludovico Sforza gave him | :09:02. | :09:05. | |
the freedom to try his hand at many different things, from architect | :09:05. | :09:13. | |
juju engineering. -- from architecture to engineering. | :09:13. | :09:18. | |
This library houses his biggest collection of mechanical drawings in | :09:18. | :09:24. | |
the world. In bold beneath the streets of Milan the Codex | :09:24. | :09:29. | |
Atlanticus is made up of 1000 sheets of his netbook. They cover a range | :09:30. | :09:34. | |
of staggering subjects, from military weapons to canal system. | :09:34. | :09:40. | |
There is even a draft of the letter Leonardo rate to Ludovico Sforza to | :09:40. | :09:46. | |
get his job. -- wrote. I wonder if you could tell | :09:46. | :09:55. | |
us what he is saying. For example, he feels he is good at building | :09:55. | :10:05. | |
:10:05. | :10:07. | ||
bridges. I can build bridges, light and strong. Does he talk about being | :10:07. | :10:13. | |
an artist as well? Music, entertainment, many other aspects of | :10:13. | :10:20. | |
his capacity. In this next drawing this is an example of the kind of | :10:20. | :10:24. | |
engineering project he took on. can see here how he is organising | :10:24. | :10:32. | |
and building the gate for different canals. | :10:32. | :10:38. | |
As well as all his engineering work, in his spare time his serious | :10:38. | :10:43. | |
interest in anatomy had been growing. A few of these anatomical | :10:43. | :10:48. | |
drawings are also tucked away here in the collection. I wonder whether | :10:48. | :10:56. | |
there is much correspondence between the mechanical drawings and the | :10:56. | :11:00. | |
anatomical studies? Sometimes he tried to combine both. Because in | :11:00. | :11:10. | |
:11:10. | :11:14. | ||
his point of view the human body is Leonardo to make his paintings of | :11:14. | :11:21. | |
the human body more effective. To start with his knowledge wasn't | :11:21. | :11:25. | |
based on first-hand observation, but on speculative classical literature | :11:25. | :11:31. | |
stretching back to Aristotle. This meant his drawings of the human body | :11:31. | :11:40. | |
were not always anatomically correct. But his enquiring mind asks | :11:40. | :11:48. | |
questions about the human form that had never been ask before. -- | :11:48. | :11:52. | |
asked. The breakthrough came at the end of the decade when he inside the | :11:52. | :12:00. | |
head of a page on a new netbook, April, 1489, in the pages that | :12:00. | :12:05. | |
follow he executed this exquisite series of drawings of the human | :12:05. | :12:10. | |
skull. They were meticulous, lucid, very precise and clearly made from | :12:10. | :12:20. | |
:12:20. | :12:23. | ||
human material gave him an enhanced understanding of anatomical | :12:23. | :12:27. | |
structures lending his drawings scientific credibility. But of | :12:27. | :12:34. | |
course it wasn't just about observation. He was also an | :12:34. | :12:38. | |
excellent draughtsman. What I'm going to do is take the skull off | :12:38. | :12:44. | |
for you, and ask you to make a series of studies of it upside down. | :12:44. | :12:48. | |
I want to find out how he articulated his understanding of the | :12:48. | :12:58. | |
body through drawing. I am back in London to meet artist Sarah Simblet. | :12:58. | :13:02. | |
What do you feel you have learned from his anatomical drawings? | :13:02. | :13:07. | |
point of reference, I have learnt technically, the use of pen and | :13:07. | :13:13. | |
ink, I have learned very much about the way in which he uses drawing to | :13:13. | :13:18. | |
see and understand structure and form, the way he uses drawing as an | :13:18. | :13:20. | |
investigative tool as well as a means of thinking and expressing | :13:20. | :13:24. | |
himself. What was his technique? | :13:24. | :13:29. | |
He has worked with a steel tipped pen, when you press it down onto the | :13:29. | :13:32. | |
paper two pieces of metal will splay apart and bite undulating the | :13:32. | :13:37. | |
pressure you can change the thickness and its oppression of your | :13:37. | :13:44. | |
line -- splay apart and by undulating the pressure. | :13:44. | :13:49. | |
He is thinking as an engineer, he is trying to understand the mechanism | :13:50. | :13:55. | |
of the body, life, whereas an awful lot of artists look at the surface | :13:55. | :13:59. | |
and want to be able to render muscular form and the power of the | :13:59. | :14:03. | |
living body, Leonardo wants to get inside and understand how it works | :14:03. | :14:06. | |
and you don't find other artists working in that way, a true | :14:06. | :14:16. | |
:14:16. | :14:18. | ||
Despite the breakthrough with the skills he put his anatomical | :14:18. | :14:25. | |
investigations aside for a decade and went on to other things. For | :14:25. | :14:28. | |
material Leonardo this meant anything from designing a game of | :14:28. | :14:34. | |
Milan's Cathedral to painting one of his master curses -- masterpieces, | :14:34. | :14:41. | |
the Last Supper. His latent enthusiasm for anatomy resurfaced | :14:41. | :14:44. | |
around 1504 and in later life the studies took up more of his time | :14:44. | :14:54. | |
:14:54. | :14:54. | ||
than any other single activity. The Royal collection's exhibition, | :14:54. | :14:57. | |
Leonardo da Vinci: The Mechanics of Man, at the Edinburgh in the, has | :14:57. | :15:02. | |
brought together a huge range of his anatomical drawings. -- Edinburgh | :15:02. | :15:07. | |
International Festival. There is one that has never been shown in the UK | :15:07. | :15:13. | |
in its entirety before. It consists of 18 sheets on which Leonardo | :15:13. | :15:19. | |
crammed more than 240 individual drawings are covering almost every | :15:19. | :15:24. | |
bone in the body, and many major muscle groups. | :15:24. | :15:30. | |
Here we see the superficial anatomy of the shoulders, and the neck. You | :15:30. | :15:37. | |
see the same model, it has been sensitively drawn, rotating in | :15:37. | :15:44. | |
space, so we get a full articulation of something which is 3-D. The | :15:44. | :15:48. | |
series continues right down to the bottom of the sheet way you can see | :15:48. | :15:52. | |
the skin has disappeared, and underneath here are the muscles in | :15:52. | :15:56. | |
the tendons laid bare so that Leonardo is not just observing how | :15:56. | :16:02. | |
things appear in one static sense before his eyes, he is always | :16:02. | :16:07. | |
thinking about how things exist in reality, in our world. He is 20 | :16:07. | :16:11. | |
articulate the functional side of anatomy. There she is trying to | :16:11. | :16:21. | |
:16:21. | :16:30. | ||
pictures to capture beautifully, a sense of physical movement. Every | :16:30. | :16:37. | |
pose has been cleverly chosen to highlight each muscle group. | :16:37. | :16:42. | |
Dancers, more than any group of people have a keen awareness of | :16:42. | :16:49. | |
their bodies and how they physically function. I have come to a rehearsal | :16:49. | :16:55. | |
of the Scottish Ballet to talk to the artistic director, Christopher | :16:55. | :17:03. | |
Hampson, about Leonardo's skilful poses in his manuscript. I find it | :17:03. | :17:07. | |
interesting he is using movement to further identify muscle groups and | :17:07. | :17:14. | |
ligaments or perhaps how far the joint will move. It is poetic he has | :17:14. | :17:20. | |
used something anatomical that could be quite dry. Christopher has found | :17:20. | :17:26. | |
a unique way of bringing his poses to life. Do you think we should | :17:26. | :17:34. | |
introduce the seminaked man? This is our principal dancer, Eric. He will | :17:34. | :17:39. | |
help us out in recreating these images. This is the first example, | :17:39. | :17:46. | |
what have you chosen? I have chosen the shoulder and the arm. It is a | :17:46. | :17:52. | |
ballet pose any way. His arm is outstretched and Eric has | :17:52. | :17:56. | |
automatically put his head looking down at the arm and you can tell | :17:56. | :18:02. | |
that is indicated. It is not snapped this way, it has a slight bend, | :18:02. | :18:06. | |
which you can see he has got there so all of the muscles are | :18:06. | :18:11. | |
well-defined. That is why I find these drawings are so interesting, | :18:11. | :18:14. | |
he has used rotation and shaping to make sure the correct the muscles | :18:14. | :18:21. | |
stand out. Why have you picked this? It has a sense of movement to | :18:21. | :18:29. | |
it. The arm is in what we call a fifth position. Eric, if you can | :18:29. | :18:35. | |
take the fifth position. This gives us the shape. By making the four arm | :18:35. | :18:41. | |
rotate in towards the head makes the bicep come out and ignite. It shows | :18:41. | :18:47. | |
the arm much more clearly. This is your third example, what is | :18:47. | :18:54. | |
happening? He is showing how the calf muscle gets fired up, ignites, | :18:54. | :19:02. | |
I presume. Then he shows this, the foot on a three quarters point. You | :19:02. | :19:09. | |
can see immediately the calf muscle gets fired up. It is a marked | :19:09. | :19:18. | |
difference. Yes.This muscle becomes hard when pulling up the heel as | :19:18. | :19:22. | |
well as releasing it. We have just seen that. You have seen it | :19:22. | :19:30. | |
perfectly. Leonardo's sketches are remarkably succinct and accurate. He | :19:30. | :19:39. | |
was able to convey all of this, simply through drawing. And now, the | :19:39. | :19:41. | |
Royal collection's Leonardo exhibition in Edinburgh is doing | :19:41. | :19:51. | |
:19:51. | :19:51. | ||
something innovative live. This is the first exhibition that compares | :19:51. | :19:54. | |
his anatomical discoveries simply using a scalpel and a pen with | :19:54. | :20:02. | |
sophisticated imaging techniques like CT and MRI scans, and also 3-D | :20:02. | :20:07. | |
films. It is staggering to reflect that even though today's anatomists | :20:07. | :20:12. | |
are using contemporary technology, many of their conclusions are | :20:12. | :20:20. | |
similar to Leonardo he made in his drawings 500 years ago. I want to | :20:20. | :20:26. | |
put this idea to the test, so I am going to do a little experiment. I | :20:26. | :20:36. | |
:20:36. | :20:38. | ||
must admit, I am feeling a little apprehensive. This is an MRI | :20:38. | :20:42. | |
scanner. Medical technology doesn't get much louder or more | :20:42. | :20:52. | |
sophisticated. Are you doing OK in there? Fine. It is going to scan my | :20:52. | :20:58. | |
hand so I can compared the results with one of Leonardo's anatomical | :20:58. | :21:05. | |
sketches that he made in the winter of 1510. OK, it is all done.I have | :21:06. | :21:13. | |
survived. You have indeed.I got through that, I still have some pins | :21:13. | :21:17. | |
and needles in my arm and left hand. It is bizarre in there it is | :21:17. | :21:25. | |
like being in a futuristic film set nightclub. I kind of went into | :21:25. | :21:31. | |
another place, another zone. I am glad it is done, I am revising -- | :21:31. | :21:37. | |
relieved. Time to get out of my gown. Talk me through what we are | :21:37. | :21:42. | |
seeing here. Even though we have all of this 21st-century technology, | :21:42. | :21:46. | |
these are still complicated images. Can you talk me through what we're | :21:46. | :21:52. | |
looking at? The straight lines are the muscles coming up. The white | :21:52. | :21:57. | |
areas of the blood vessels. We have the bones over here which are dark | :21:57. | :22:03. | |
on this picture. I brought along a reproduction of one of the most | :22:03. | :22:08. | |
famous sheets by Leonardo. This was done in the winter of 1510, and it | :22:08. | :22:14. | |
is startling looking at it compared to these images. Absolutely | :22:14. | :22:21. | |
accurate. All of these tendons radiating up from the rest. It is | :22:21. | :22:26. | |
amazing how accurate he was able to draw it. We can see the different | :22:26. | :22:32. | |
bones, he has done it absolutely correct. This is a knot of | :22:32. | :22:37. | |
complexity isn't it? Yes, these are the bones in the wrist which allow | :22:37. | :22:44. | |
us to do everything. It looks like an area of parched earth. What is | :22:44. | :22:48. | |
brilliant about these drawings is he manages to take something that is | :22:49. | :22:52. | |
clearly so complicated and make it lucid and clear in a sheet done 500 | :22:52. | :23:01. | |
years ago. To achieve this level of accuracy, Leonardo not only had to | :23:01. | :23:05. | |
be an excellent draughtsman, but also he had to be handy with a | :23:05. | :23:12. | |
scalpel and have direct access to human bodies. In the course of his | :23:12. | :23:17. | |
anatomical investigations, he only dissected about 30 corpses. And | :23:17. | :23:24. | |
around 20 of these were carried out while he was compiling his | :23:24. | :23:32. | |
manuscript and was probably collaborating with a doctor at the | :23:32. | :23:37. | |
University macro. I have come to the anatomy department at Glasgow | :23:37. | :23:42. | |
University to see the human dissection. What I hope is that this | :23:42. | :23:47. | |
will help me understand Leonardo's achievement in these drawings by | :23:47. | :23:55. | |
witnessing the complexity of what he himself would have observed. We are | :23:55. | :23:59. | |
going to try and imitate one of the famous drawings Leonardo did, and | :23:59. | :24:04. | |
give you an idea how complex the tissue is and how good a job he | :24:04. | :24:12. | |
actually did. The difference between the modern era and his time is he | :24:12. | :24:17. | |
obtained bodies either illegally through the church and through the | :24:17. | :24:24. | |
pauper's house. But today, everyone we use here is donated willingly and | :24:24. | :24:27. | |
knowingly. In their actual life, they have signed the forms | :24:27. | :24:34. | |
themselves. We will move to the foot, and we will see the tendons | :24:34. | :24:38. | |
becoming more prominent. These are the things standing out in his | :24:38. | :24:42. | |
drawings. They are much more pronounced in his drawings and in | :24:42. | :24:47. | |
reality. If you did not know what you are looking for, you might miss | :24:47. | :24:52. | |
it? It is testament to the scale of Leonardo he can take something and | :24:52. | :25:01. | |
make it so clear. You can see underneath, each tendon which is the | :25:01. | :25:09. | |
thick white band? One of each of these for each toe. A collection of | :25:09. | :25:14. | |
strings or ropes connected to the muscle at one end and to the bone at | :25:14. | :25:20. | |
the other. Remarkable that not only he captured them in terms of his | :25:20. | :25:25. | |
drawings, but understood the mechanical purpose of them. I | :25:25. | :25:31. | |
imagine Leonardo going through this dissection and not just looking at | :25:31. | :25:34. | |
it as a static example, but pulling on the tendons and moving things | :25:34. | :25:41. | |
around. It is so clear to me now he did something remarkable in the | :25:41. | :25:51. | |
:25:51. | :25:52. | ||
drawings to articulate that in a clever, simple, plain fashion. | :25:52. | :25:57. | |
Leonardo's combined skills as a dissected and draughtsman, meant he | :25:57. | :26:00. | |
would glean insight that would not be observed to gain for hundreds of | :26:00. | :26:07. | |
years. The full scope of his scientific accomplishments can be | :26:07. | :26:10. | |
seen in the field of cardiac anatomy, which he carried out | :26:10. | :26:18. | |
towards the end of his career. Intrigued by the way the aortic | :26:18. | :26:23. | |
valve opens and closes to ensure blood flows in one direction, | :26:23. | :26:28. | |
Leonardo set about constructing a model, filling an ox's heart with | :26:28. | :26:35. | |
wax. When the wax had hardened, he recreated the structure in glass and | :26:35. | :26:41. | |
pumped a mixture of grass seeds, suspended in water, through it. It | :26:41. | :26:48. | |
allowed him to observe the vortex of the seed swelling around at the base | :26:48. | :26:55. | |
of the aorta. And the result, Leonardo correctly posited, it | :26:55. | :26:59. | |
helped to close the aortic valve. And that would not be observed a | :27:00. | :27:07. | |
game for hundreds of years, until the 20th century. -- a game. One | :27:07. | :27:12. | |
contemporary practitioner who studied this side of Leonardo's work | :27:12. | :27:17. | |
is the heart surgeon, Francis Wells. The first time I focused on the | :27:17. | :27:22. | |
drawings was when I'd just qualified as a doctor. Once I had seen them, I | :27:22. | :27:26. | |
thought they were far better than anything we had in the current day, | :27:26. | :27:33. | |
modern textbook of anatomy. They were beautiful, accurate and | :27:33. | :27:37. | |
absorbing. You can look with a magnifying glass at some of the | :27:37. | :27:43. | |
heart for example and the fineness of the shading gives you a depth to | :27:43. | :27:50. | |
these drawings, which is technical genius. What did he find out about | :27:50. | :27:57. | |
the heart? The heart was thought of as a two chamber structure up until | :27:57. | :28:02. | |
and after his time - of course he never published this. But the | :28:02. | :28:07. | |
atria, the filling changes were part as the system, the heart. Leonardo | :28:08. | :28:13. | |
firmly states the heart is for Chambers. I brought along a | :28:13. | :28:20. | |
reconstruction of a drawing he made where he observes the vortices. | :28:20. | :28:28. | |
is a fine example of synopsis. He has the description of the vortices, | :28:28. | :28:32. | |
and little diagrams showing how this argument has to be the right one for | :28:32. | :28:37. | |
the mechanism of closure for the valve, and not such simple reflex of | :28:37. | :28:42. | |
the blood and how it would fail. This wasn't understood or known | :28:42. | :28:48. | |
about until the last century. But it was reported in 1968 by two | :28:48. | :28:55. | |
engineers in Oxford. The references were back to Leonardo da Vinci, 500 | :28:55. | :29:02. | |
years old. The tragedy of his anatomical investigations is that he | :29:03. | :29:07. | |
never got round to publishing them. It was almost as if he was | :29:07. | :29:11. | |
constantly getting sidetracked with all of his difference projects. The | :29:11. | :29:20. | |
abrupt death of his collaborator from the plague in 1511, coupled | :29:20. | :29:25. | |
with public -- political turmoil in Milan cut short his systematic | :29:25. | :29:33. | |
efforts to finish his treaties. When he died in 1519, the hundreds of | :29:33. | :29:37. | |
sheets and notes he compiled over three decades remained hidden among | :29:37. | :29:44. | |
his private papers. If Leonardo had published his treaties on anatomy, E | :29:44. | :29:51. |